After being away from his city for a long time Toni Cipriani returns to Liberty City (the most dangerous city in America) to continue to work with his mob boss, Salvatore Leone. On his return he discovers that not everything is as before: in fact the order of the boss will have to follow the orders of Vincenzo Cilli, her manager. In addition to Vincent know JD O'Toole, owner of a club Sindacco (Lion of the rival family), Maria Latore But the woman's boss and the mother of Toni Cipriani. The relationship breaks down when Vincenzo Cipriani instructs to fetch his car that was actually a police trap. The two will clash in a ship in Portland where Toni kills Vincenzo. Killed Vincent, Salvatore and Toni will work for both will be required to reach Staunton Island to understand the intentions of the other gang Mafia. Here will know that Toni Donald Love, a multimillionaire who aspires to the office of mayor of Liberty City and Leon Mc Affrey a corrupt cop. Salvatore meanwhile, not knowing how and why it ends up in prison Shorside Vale, last island of Liberty City. Meanwhile, Toni knows Toshiko Kasen wife of the head of the Yakuza family (who later kill). After killing Paulie Sindacco, Salvatore is released along with Toni and Massimo kill Turin and all who conspired against the family plans Leone.
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